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Article: A new peculiar dinosaur egg, Sankofa pyrenaica oogen. nov. oosp. nov. from the Upper Cretaceous coastal deposits of the Aren Formation, south-central Pyrenees, Lleida, Catalonia, Spain

Publication: Palaeontology
Volume: 55
Part: 2
Publication Date: March 2012
Page(s): 325 339
Author(s): <p>Nieves L&oacute;pez-Mart&iacute;nez and Enric Vicens</p>
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How to Cite

LÓPEZ-MART͍NEZ, N., VICENS, E. 2012. A new peculiar dinosaur egg, Sankofa pyrenaica oogen. nov. oosp. nov. from the Upper Cretaceous coastal deposits of the Aren Formation, south-central Pyrenees, Lleida, Catalonia, Spain. Palaeontology55, 2, 325–339.

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  • ARDÈVOL, L., KLIMOWITZ, J., MALAGÓN, J. and NAGTEGAAL, P. J. C. 2000. Depositional sequence response to foreland deformation in the upper Cretaceous of the Southern Pyrenees, Spain. Bulletin of the American Association of Petroleum Geology, 84, 566–587.
  • BACETA, J. I., PUJALTE, V. and CABALLERO, F. 2006. Post conference fieldtrip guidebook. Climate and Biota of the Early Paleogene, Bilbao.
  • BARTA, Z. and SZEKELY, T. 1997. The optimal shape of avian eggs. Functional Ecology, 11, 656–662.
  • BRAVO, A. M., MORATALLA, J. J., SANTAFÉ, J. V. and DE SANTISTEBAN, C. 2000. Faidella, a new upper Cretaceous nesting site from the Tremp Basin (Lérida province, Spain). 15–21. In BRAVO, A. M. and REYES, T. (eds). First International Symposium on dinosaur eggs and babies. Extended abstracts. Publicacions de la Diputació de Lleida, Lleida, 192 pp.
  • CARPENTER, K. 1999. Eggs, nests, and baby dinosaurs: a look at dinosaur reproduction. Indiana University Press, Bloomington, 336 pp.
  • CHIAPPE, L. M., CORIA, A. C., DINGUS, L., JACKSON, F., CHINASAMY, A. and FOX, M. 1998. Sauropod dinosaur embryos from the Late Cretaceous of Patagonia. Nature, 396, 258–261.
  • LARK, J. M., NORELL, M. A. and CHIAPPE, L. M. 1999. An oviraptorid skeleton from the Late Cretaceous of Ukhaa Tolgod, Mongolia, preserved in an avianlike brooding position over an oviraptorid nest. American Museum Novitates, 3265, 1–36.
  • DÍAZ-MOLINA, M. 1987. Sedimentación sintectónica asociada a una subida relativa del nivel del mar durante el Cretácico superior (Fm. Tremp, provincia de Lérida). 69–93. In SANZ, J. L. (ed.). Geología y Paleontología (Arcosaurios) de los yacimientos de Galve (Teruel, Cretácico inferior) y Tremp (Lérida, Cretácico superior). Estudios Geológicos, Volumen extraordinario Galve-Tremp, Madrid, 110 pp.
  • DÍAZ-MOLINA, M., KÄLIN, O., BENITO, M. I., LOPEZ-MARTINEZ, N. and VICENS, E. 2007. The depositional setting of the dinosaur eggshell-bearing Aren Fm at Bastus (Late Cretaceous, South-Central Pyrenees). Sedimentary Geology, 199, 205–221.
  • DONAIRE, M. and LOPEZ-MARTINEZ, N. 2009. Porosity of Late Paleocene Ornitholithus eggshells (Tremp Fm, south-central Pyrenees, Spain): palaeoclimatic implications. Palaeogeography, Palaeoclimatology, Palaeoecology, 279, 147–159.
  • ESCUER, J., DELCLÒS, X., COLOMBO, F., BRAVO, A., PERALBA, J., MARTÍ, X., ORTEGA, J., MIQUELA, R. and PARELLADA, C. 2006. General distribution of dinosaur eggshells and clutches in the upper Cretaceous of Coll de Nargó area (South-central Pyrenees, Spain). Ameguiniana, 44(Suppl. 4), 17R.
  • GARCIA, G. 1998. Les coquilles d’oeufs de dinosaures du Crétacé supérieur du Sud de la France: Diversité, paléobiologie, biochronologíe et paléoenvironments. PhD Thesis, Vol. 2. Académie de Montpellier, 152 pp.
  • GARCIA, G., MARIVAUX, L., PÉLISSIÉ, T. and VIANEY-LIAUD, M. 2006. Earliest Laurasian sauropod eggshells. Acta Palaeontologica Polonica, 51, 99–104.
  • GRELLET-TINNER, G. and CHIAPPE, L. M. 2004. Dinosaur eggs and nesting: implications for understanding the origin of birds. 185–214. In CURRIE, P. J., KOPPELHUS, E. B., SHUGAR, M. A. and WRIGHT, J. L. (eds). Feathered dragons: studies on the transition from dinosaurs to birds. Indiana University Press, Bloomington, 268 pp.
  • GRELLET-TINNER, G. and MAKOVICKY, P. 2006. A possible egg of the dromaeosaur Deinonychus antirrhopus: phylogenetic and biological implications. Canadian Journal of Earth Sciences, 43, 705–719.
  • GRELLET-TINNER, G., CHIAPPE, L. M., NORELL, M. and BOTTJER, D. 2006. Dinosaur eggs and nesting behaviors: a paleobiological investigation. Palaeogeography, Palaeoclimatology, Palaeoecology, 232, 294–321.
  • HORNER, J. 2000. Dinosaur reproduction and parenting. Annual Review Earth and Planetary Sciences, 28, 19–45.
  • LÓPEZ-MARTÍNEZ, N. 2000. Eggshell sites from the Cretaceous-Tertiary transition in South-central Pyrenees (Spain). 95–115. In BRAVO, A. and REYES, T. (eds). First International Symposium on dinosaur eggs and babies. Extended abstracts. Publicacions de la Diputació de Lleida, Lleida, 192 pp.
  • LÓPEZ-MARTÍNEZ, N., MORATALLA, J. J. and SANZ, J. L. 2000. Dinosaur nesting in tidal flats. Palaeogeography, Paleoclimatology, Paleoecology, 160, 153–163.
  • LÓPEZ-MARTÍNEZ, N., ARDÉVOL, L., ARRIBAS, M. E., CANUDO, J. I., LACASA, A., PEREDA-SUBERBIOLA, X., SOLER-GIJÓN, R., TORICES, A. and VICENS, E. 2002. The record of the dinosaur extinction in the South-Central Pyrenees. First International Paleontological Congress, Macquarie University, Australia 6–10 July 2002, Abstracts p. 234.
  • LÓPEZ-MARTÍNEZ, N., GALOBART, À. and VICENS, E. (coord.) 2009. Upper Cretaceous continental deposits of South-eastern and South-central Pyrenees. Fundamental! Teruel, 14, 1–91.
  • LOYAL, R. S., MOHABEY, D. M., KHOSLA, A. and SHANI, A. 1998. Status and palaeobiology of the Late Cretaceous Indian Theropods with the description of a new eggshell oogenus and oospecies, Ellipsoolithus khedaensis from the Lameta Formation, district Kheda, Gujarat, Western India. Gaia (Lisbon), 15, 379–387.
  • MIKHAILOV, K. E. 1992. The microstructure of avian and dinosaurian eggshell: phylogenetic implications. In CAMPBELL, K. E. JR (ed.). Papers in avian paleontology Honoring Pierce Brodkorb. Natural History Museum of Los Angeles County, Science Series, 36, 361–373.
  • MIKHAILOV, K. E. 1997. Fossil and recent eggshell in amniotic vertebrates: fine structure comparative morphology and classification. Special Papers in Palaeontology, 56, 1–80.
  • OMS, O. and CANUDO, J. I. 2004. Datación magnetoestratigráfica de los dinosaurios del Cretácico terminal (Maastrichtiense superior) de Arén (Huesca, Unidad Surpirenaica Central). Geo-Temas, 6, 51–54.
  • OMS, O., DINARÉS-TURELL, J., VICENS, E., ESTRADA, R., VILA, B., GALOBART, À. and BRAVO, A. M. 2007. Integrated stratigraphy from the Vallcebre Basin (southeastern Pyrenees, Spain): new insights on the continental Cretaceous-Tertiary transition in southwest Europe. Palaeogeography, Palaeoclimatology, Palaeoecology, 255, 35–47.
  • PEREDA-SUBERBIOLA, X., CANUDO, J. J., CRUZADO-CABALLERO, P., BARCO, J. L., LÓPEZ-MARTÍNEZ, N., OMS, O. and RUIZ-OMEÑACA, J. I. 2009. The last hadrosaurid dinosaurs of Europe: a new lambeosaurine from the uppermost Cretaceous of Arén (Huesca, Spain). Comptes Rendues Paleovol, 8, 559–572.
  • SANZ, J. L., MORATALLA, J., DIAZ-MOLINA, M., LÓPEZ-MARTÍNEZ, N., KÄLIN, O. and VIANEY-LIAUD, M. 1995. Dinosaur nest at the sea shore. Nature, 376, 731–732.
  • SCHWEITZER, M. H., JACKSON, F. D., CHIAPPE, L. M., SCHMITT, J. G., CALVO, J. O. and RUBILAR, D. E. 2002. Late Cretaceous avian eggs with embryos from Argentina. Journal of Vertebrate Paleontology, 22, 191–195.
  • SMART, I. H. M. 1995. Egg shape in birds. 101–117. In DEEMING, C. D. and FERGUSON, M. W. J. (eds). Egg incubation: its effects on embryonic development. Cambridge University Press, New York, 250 pp.
  • TORICES, A. 2007. Los dinosaurios terópodos del Cretácico superior de la Cuenca Sur-Pirenaica. Unpublished PhD Thesis. Universidad Complutense de Madrid, Madrid, 409 pp.
  • VARRICCHIO, D. J. and JACKSON, F. D. 2004. A phylogenetic assessment of prismatic dinosaur eggs from the Cretaceous Two Medicine Formation of Montana. Journal of Vertebrate Paleontology, 24, 931–937.
  • VARRICCHIO, D. J., JACKSON, F., BORKOWSKI, J. and HORNER, J. R. 1997. Nest and egg clutches of the dinosaur Troodon formosus and the evolution of avian reproductive traits. Nature, 385, 247–250.
  • VARRICCHIO, D. J., HORNER, J. R. and JACKSON, F. 2002. Embryos and eggs for the Cretaceous theropod dinosaur Troodon formosus. Journal of Vertebrate Paleontology, 22, 564–576.
  • VIANEY-LIAUD, M. and LÓPEZ-MARTÍNEZ, N. 1997. Late Cretaceous dinosaur eggshells from the Tremp basin (Southern Pyrenees, Lleida, Spain). Journal of Paleontology, 71, 1157–1171.
  • VIANEY-LIAUD, M., MALLAN, P., BUSCAIL, O. and MONTGELARD, C. 1994. Review of French dinosaur eggshells: morphology, structure, mineral, and organic composition. 151–183. In CARPENTER, K., HIRSCH, K. and HORNER, J. R. (eds). Dinosaur eggs and babies. Cambridge University Press, Cambridge, 376 pp.
  • VICENS, E., ARDÈVOL, L. and LÓPEZ-MARTÍNEZ, N. 1999. The Rudists as biostratigraphic markers: an example from the Late Cretaceous of the Southern Pyrenees, Spain. Erlanger Geologische Abhandlungen, 3, 74–75.
  • VICENS, E., ARDÈVOL, L., LÓPEZ-MARTÍNEZ, N. and ARRIBAS, M. A. 2004. Rudist biostratigraphy in the upper Cretaceous of the Southern Pyrenees, Spain. Courier Seckenberg, Frankfurt, 247, 113–127.
  • ZELENITSKY, D. K. and HILLS, L. V. 1996. An egg clutch of Prismatoolithus levis oosp. nov. from the Oldman Formation (Upper Cretaceous), Devil’s Coulee, southern Alberta. Canadian Journal of Earth Sciences, 33, 1127–1131.
  • ZELENITSKY, D. K. and THERRIEN, F. 2008a. Phylogenetic analysis of reproductive traits of maniraptoran theropods and its implications for egg parataxonomy. Palaeontology, 51, 807–816.
  • ZELENITSKY, D. K. and THERRIEN, F. 2008b. Unique maniraptoran egg clutch from the upper Cretaceous Two Medicine Formation of Montana reveals theropod nesting behaviour. Palaeontology, 51, 1253–1259.
  • ZELENITSKY, D. K., MODESTO, S. P. and CURRIE, P. J. 2002. Bird-like characteristics of troodontid theropod eggshell. Cretaceous Research, 23, 297–305.
  • ZHAO, Z.-K. 1975. The microstructure of dinosaurian eggshells of Nanhsiung, Kwangtung and Guangdong. Vertebrata PalAsiatica, 13, 105–117.
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