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Article: A new skeleton of the therocephalian synapsid Olivierosuchus parringtoni from the Lower Triassic South African Karoo Basin

Publication: Palaeontology
Volume: 54
Part: 3
Publication Date: May 2011
Page(s): 591 606
Author(s): Jennifer Botha-Brink and Sean P. Modesto
Addition Information

How to Cite

BOTHA-BRINK, J., MODESTO, S. P. 2011. A new skeleton of the therocephalian synapsid Olivierosuchus parringtoni from the Lower Triassic South African Karoo Basin. Palaeontology54, 3, 591–606.

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  • ABDALA, F. 2007. Redescription of Platycraniellus elegans (Therapsida, Cynodontia) from the Lower Triassic of South Africa, and the cladistic relationships of eutheriodonts. Palaeontology, 50, 591–618.
  • ABDALA, F., RUBIDGE, B. S. and HEEVER van den, J. 2008. The oldest therocephalians (Therapsida, Eutheriodontia) and the early diversification of Therapsida. Palaeontology, 51, 1011–1024.
  • ANDERSON, J. M. 1968. The confused state of classification within the family Procynosuchidae. Palaeontologia Africana, 11, 77–83.
  • ATTRIDGE, J. 1956. The morphology and relationships of a complete therocephalian skeleton from the Cistecephalus Zone of South Africa. Proceedings of the Royal Society, Edinburgh B, IV, 59–93.
  • BARRY, T. H. 1965. On the epipterygoid-alisphenoid transition in Therapsida. Annals of the South African Museum, 48, 399–426.
  • BOONSTRA, L. D. 1934. A contribution to the morphology of the mammal-like reptiles of the suborder Therocephalia. Annals of the South African Museum, 31, 215–267.
  • BOONSTRA, L. D. 1954. Scaloposaurid from the Tapinocephalus Zone. Annals and Magazine of Natural History, 7, 154–165.
  • BOONSTRA, L. D. 1964. The girdles and limbs of the pristerognathid Therocephalia. Annals of the South African Museum, 48, 121–165.
  • BOTHA, J. and SMITH, R. M. H. 2006. Rapid vertebrate recuperation in the Karoo Basin of South Africa following the End-Permian extinction. Journal of African Earth Sciences, 45, 502–514.
  • BOTHA, J. and SMITH, R. M. H. 2007. Lystrosaurus species composition across the Permo-Triassic boundary of South Africa. Lethaia, 40, 125–137.
  • BOTHA, J., ABDALA, F. and SMITH, R. M. H. 2007. The oldest cynodont: new clues on the origin and early diversification of the Cynodontia. Zoological Journal of the Linnean Society, 149, 447–492.
  • BREMER, K. 1988. The limits of amino acid sequence data in angiosperm phylogenetic reconstruction. Evolution, 42, 795–803.
  • BRINK, A. S. 1956. On Aneugomphius ictidoceps Broom and Robinson. Palaeontologia Africana, 4, 97–115.
  • BRINK, A. S. 1957. On the skeleton of Aneugomphius ictidoceps Broom and Robinson. Palaeontologia Africana, 5, 29–37.
  • BRINK, A. S. 1958. Notes on some whaitsiids and moschorhinids. Palaeontologia Africana, 6, 23–49.
  • BRINK, A. S. 1960. On some small therocephalians. Palaeontologia Africana, 7, 155–182.
  • BRINK, A. S. 1963. On Bauria cynops Broom. Palaeontologia Africana, 8, 39–56.
  • BRINK, A. S. 1965. A new ictidosuchid (Scaloposauria) from the Lystrosaurus-Zone. Palaeontologia Africana, 9, 129–138.
  • BRINK, A. S. 1986. Illustrated bibliographic catalogue of the Synapsida. Parts 1 and 2. Department of Mineral and Energy Affairs, Pretoria, Handbook 10, 540 pp.
  • BROOM, R. 1901. On Ictidosuchops primaevus. Transactions of the South African Philosophical Society, 11, 177–184.
  • BROOM, R. 1903. On the classification of the theriodonts and their allies. Report of the South African Association for the Advancement of Science, 1, 286–294.
  • BROOM, R. 1906. On the Permian and Triassic faunas of South Africa. Geological Magazine of New Series, 5, 36.
  • BROOM, R. 1936. On some new genera and species of Karroo fossil reptiles, with notes on some others. Annals of the Transvaal Museum, 18, 349–386.
  • BROOM, R. 1938. On a nearly complete therocephalian skeleton. Annals of the Transvaal Museum, 19, 257–261.
  • BROOM, R. 1948. The skeleton of a very small therocephalian. Annals of the Transvaal Museum, 21, 39–41.
  • CATUNEANU, O., WOPFNER, H., ERIKSSON, P. G., CAIRNCROSS, B., RUBIDGE, B. S., SMITH, R. M. H. and HANCOX, P. J. 2005. The Karoo basins of south-central Africa. Journal of African Earth Sciences, 43, 211–253.
  • CLUVER, M. A. 1969. Zorillodontops, a new scaloposaurid from the Karoo. Annals of the South African Museum, 52, 183–188.
  • COLBERT, E. H. 1948. The mammal-like reptile Lycaenops. Bulletin of the American Museum of Natural History, 89, 353–404.
  • COLBERT, E. H. and KITCHING, J. W. 1981. Scaloposaurian reptiles from the Triassic of Antarctica. American Museum Novitates, 2709, 8–16.
  • CYS, J. M. 1967. Osteology of the pristerognathid Cynariognathus platyrhinus (Reptilia: Theriodontia). Journal of Paleontology, 41, 776–790.
  • DURAND, J. F. 1991. A revised description of the skull of Moschorhinus (Therapsida, Therocephalia). Annals of the South African Museum, 99, 381–413.
  • FINDLAY, G. H. 1968. On the scaloposaurid skull of Olivieria parringtoni, Brink with a note on the origin of hair. Palaeontologia Africana, 11, 47–59.
  • FOURIE, H. 2001. Morphology and function of the postcrania of selected genera of Therocephalia (Amniota: Therapsida). PhD dissertation, University of the Witwatersrand, Johannesburg, 185 pp.
  • FOURIE, H. and RUBIDGE, B. S. 2007. The postcranial skeletal anatomy of the therocephalian Regisaurus (Therapsida: Regisauridae) and its utilization for biostratigraphic correlation. Palaeontologia Africana, 42, 1–16.
  • GROENEWALD, G. H. and KITCHING, J. W. 1995. Biostratigraphy of the Lystrosaurus Assemblage Zone. 35–39. In RUBIDGE, B. S. (ed.). Biostratigraphy of the Beaufort Group (Karoo Supergroup). Council for Geoscience, Pretoria, 46 pp.
  • HEEVER, J. A. V A N D E N. 1994. The cranial anatomy of the early Therocephalia. Annale van die Universiteit van Stellenbosch, 1, 1–59.
  • HOPSON, J. A. and BARGHUSEN, H. R. 1986. An analysis of therapsid relationships. 83–106. In HOTTON, N.III, MACLEAN, P. D., ROTH, J. J. and ROTH, E. C. (eds). The ecology and biology of mammal-like reptiles. Smithsonian Institution Press, Washington, 326 pp.
  • HOU, L. 1979. On a new theriodont from Inner Mongolia. Vertebrata PalAsiatica, 17, 1–130.
  • HUTTENLOCKER, A. 2009. An investigation into the cladistic relationships and monophyly of therocephalian therapsids (Amniota: Synapsida). Zoological Journal of the Linnean Society, 157, 865–891.
  • KAMMERER, C. 2008. A new therocephalian from the Cistecephalus Assemblage Zone of South Africa and new information on therocephalian systematics. Journal of Vertebrate Paleontology, 28(Suppl. 3), 98A–99A.
  • KAMMERER, C. and SIDOR, C. A. 2002. Replacement names for the therapsid genera Criocephalus Broom 1928 and Olivieria Brink 1965. Palaeontologia Africana, 38, 71–72.
  • KEMP, T. S. 1972. Whaitsiid Therocephalia and the origin of cynodonts. Philosophical Transactions of the Royal Society of London, Series B, 264, 1–54.
  • KEMP, T. S. 1978. Stance and gait in the hindlimb of a therocephalian mammal-like reptile. Journal of the Zoological Society of London, 186, 143–161.
  • KEMP, T. S. 1979. The primitive cynodont Procynosuchus: functional anatomy of the skull and relationships. Philosophical Transactions of the Royal Society of London, Series B, 285, 73–122.
  • KEMP, T. S. 1980. The primitive cynodont Procynosuchus: structure, function, and the evolution of thepostcranial skeleton. Philosophical Transactions of the Royal Society of London, Series B, 288, 217–258.
  • KEMP, T. S. 1986. The skeleton of a baurioid therocephalian therapsid from the Lower Triassic (Lystrosaurus Zone) of South Africa. Journal of Vertebrate Paleontology, 6, 215–232.
  • KING, G. M. 1996. A description of the skeleton of a bauriid therocephalian from the Early Triassic of South Africa. Annals of the South African Museum, 104, 379–393.
  • KITCHING, J. W. 1970. A short review of the Beaufort zoning in South Africa. 309–312. In HAUGHTON, S. H. (ed.). Second Symposium on Gondwana Stratigraphy and Palaeontology. Council for Scientific and Industrial Research, Pretoria, 689 pp.
  • MENDREZ, C. H. 1972. On the skull of Regisaurus jacobi, a new genus and species of Bauriamorpha Watson and Romer 1956 (= Scaloposauria Boonstra 1953), from the Lystrosaurus-zone of South Africa. 191–212. In JOYSEY, K. and KEMP, T. S. (eds). Studies in vertebrate evolution. Oliver and Boyd, Edinburgh, 284 pp.
  • MENDREZ, C. H. 1974. A new specimen of Promoschorhynchus platyrhinus Brink 1954 (Moschorhinidae) from the Daptocephalus-Zone (Upper Permian) of South Africa. Palaeontologia Africana, 17, 69–85.
  • MENDREZ, C. H. 1975. Principales variations du palais chez les Thérocéphales Sud-Africains (Pristerosauria et Scaloposauria) au cours du Permien supérieur et du Trias inférieur. Problèmes Actueles de Paléontologie-Évolution de Vertébrés, Colloque International C.N.R.S., 218, 379–408.
  • NEVELING, J. 2004. Stratigraphic and sedimentological investigation of the contact between the Lystrosaurus and the Cynognathus Assemblage Zones (Beaufort Group: Karoo Supergroup). Bulletin of the Council for Geoscience, 137, 1–165.
  • NEVELING, J., HANCOX, P. J. and RUBIDGE, B. S. 2006. Biostratigraphy of the lower Burgersdorp Formation (Beaufort Group; Karoo Supergroup) of South Africa – implications for the stratigraphic ranges for Early Triassic tetrapods. Palaentologia Africana, 48, 81–87.
  • NOPCSA, F. 1923. Die Familien der Reptilien. Verlag von Gebrüder Borntraeger, Berlin, 210 pp.
  • OSBORN, H. F. 1903. On the primary division of the Reptilia into two sub-classes, Synapsida and Diapsida. Science, 17, 275–276.
  • RETALLACK, G. J., SMITH, R. M. H. and WARD, P. D. 2003. Vertebrate extinction across Permian-Triassic boundary in Karoo Basin, South Africa. GSA Bulletin, 115, 1133.
  • ROMER, A. S. 1956. Osteology of reptiles. University of Chicago Press, Chicago, 772 pp.
  • ROWE, T. 1988. Definition, diagnosis, and origin of Mammalia. Journal of Vertebrate Paleontology, 8, 241–264.
  • RUBIDGE, B. S. (ed.) 1995. Biostratigraphy of the Beaufort Group (Karoo Supergroup). Geological Survey of South Africa, Pretoria, 46 pp.
  • RUBIDGE, B. S. and SIDOR, C. A. 2001. Evolutionary patterns among Permo-Triassic therapsids. Annual Review of Ecology and Systematics, 32, 449–480.
  • SIGOGNEAU, D. 1963. Note sur une nouvelle espèce de Scaloposauridae. Palaeontologia Africana, 8, 13–37.
  • SIGOGNEAU, D. 1970. Révision systematique des gorgonopsiens sud-Africains. Cahiers de Paléontologie, 1970, 1–416.
  • SMITH, R. M. H. and BOTHA, J. 2005. The recovery of terrestrial vertebrate diversity in the South African Karoo Basin after the End-Permian extinction. Comptes Rendus Palevol, 4, 555–568.
  • SMITH, R. M. H. and WARD, P. D. 2001. Pattern of vertebrate extinctions across an event bed at the Permian-Triassic boundary in the Karoo Basin of South Africa. Geology, 29, 1147–1150.
  • VICKARYOUS, M. K. and HALL, B. K. 2006. Homology of the reptilian coracoid and a reappraisal of the evolution and development of the amniote pectoral apparatus. Journal of Anatomy, 208, 263–285.
  • WARD, P. D., MONTGOMERY, D. R. and SMITH, R. 2000. Altered river morphology in South Africa related to the Permian-Triassic Extinction. Science, 289, 1741–1743.
  • WARD, P. D., BOTHA, J., BUICK, R., KOCK de, M. O., ERWIN, D. H., GARRISON, G., KIRSCHVINK, J. and SMITH, R. 2005. Abrupt and gradual extinction among Permian land vertebrates in the Karoo Basin, South Africa. Science, 307, 709–713.
  • WATSON, D. M. S. 1931. On the skeleton of a bauriamorph reptile. Proceedings of the Zoological Society of London, 1931, 1163–1205.
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