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Article: Testing for a decline in diversity prior to extinction: Languedocian (latest mid-Cambrian) distribution of cinctans (Echinodermata) in the Iberian Chains, NE Spain

Publication: Palaeontology
Volume: 53
Part: 6
Publication Date: November 2010
Page(s): 1349 1368
Author(s): <p>Samuel Zamora and J. Javier &Aacute;lvaro</p>
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ZAMORA, S., ÁLVARO, J. 2010. Testing for a decline in diversity prior to extinction: Languedocian (latest mid-Cambrian) distribution of cinctans (Echinodermata) in the Iberian Chains, NE Spain. Palaeontology53, 6, 1349–1368.

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  • ÁLVARO, J. J. and CLAUSEN, S. 2005. Major geodynamic and sedimentary constraints on the chronostratigraphic correlation of the Lower–Middle Cambrian transition in the western Mediterranean region. Geosciences Journal, 9, 145–160.
  • ÁLVARO, J. J. and VENNIN, E. 1996. The Middle–Upper Cambrian transition in the Iberian Chains (NE Spain). An integrated approach. Rivista Italiana di Paleontologia e Stratigrafia, 102, 341–352.
  • ÁLVARO, J. J. and VENNIN, E. 1997. Episodic development of Cambrian eocrinoid-sponge meadows in the Iberian Chains (NE Spain). Facies, 37, 49–64.
  • ÁLVARO, J. J. and VENNIN, E. 2001. Benthic marine communities recorded in the Cambrian Iberian platform, NE Spain. Palaeontographica Abteilung A, 262, 1–23.
  • ÁLVARO, J. J. and VIZCAÏNO, D. 1998. Révision biostratigraphique du Cambrien moyen du versant méridional de la Montagne Noire (Languedoc, France). Bulletin de la Société Géologique de France, 169, 233–242.
  • ÁLVARO, J. J. and VIZCAÏNO, D. 2001. Evolutionary trends in the ornamentation of Cambrian solenopleuropsine trilobites. Palaeontology, 44, 131–141.
  • ÁLVARO, J. J. and VIZCAÏNO, D. 2003. The conocoryphid biofacies, a benthic assemblage of normal-eyed and blind trilobites. Special Papers in Palaeontology, 70, 127–140.
  • ÁLVARO, J. J., VIZCAÏNO, D. and VENNIN, E. 1999. Trilobite diversity patterns in the Middle Cambrian of southwestern Europe: a comparative study. Palaeogeography, Palaeoclimatology, Palaeoecology, 151, 241–254.
  • ÁLVARO, J. J., VENNIN, E., MUÑOZ, A., SÁNCHEZ-VALVERDE, B. and OJEDA, J. L. 2000. Interplay of orbital forcing and tectonic pulses in the Cambrian Iberian platform, NE Spain. International Journal of Earth Sciences, 89, 366–376.
  • ÁLVARO, J. J., LEFEBVRE, B., SHERGOLD, J. H. and VIZCAÏNO, D. 2001. The Middle–Upper Cambrian of the southern Montagne Noire. Annales de la Société Géologique du Nord (2e Série), 8, 205–211.
  • ÁLVARO, J. J., GONZÁLEZ-GÓMEZ, C. and VIZCAÏNO, D. 2003a. Paleogeographic patterns of the Cambrian-Ordovician transition in the southern Montagne Noire (France): preliminary results. Bulletin de la Société Géologique de France, 174, 23–31.
  • ÁLVARO, J. J., ELICKI, O., GEYER, G., RUSHTON, A. W. A. and SHERGOLD, J. H. 2003b. Palaeogeographical controls on the Cambrian trilobite immigration and evolutionary patterns reported in the western Gondwana margin. Palaeogeography, Palaeoclimatology, Palaeoecology, 195, 5–35.
  • ÁLVARO, J. J., VIZCAïNO, D., KORDULE, V., FATKA, O. and PILLOLA, G. L. 2004. Some solenopleurine trilobites from the Languedocian (late Mid Cambrian) of western Europe. Geobios, 37, 135–147.
  • ÁLVARO, J. J., FERRETTI, F., GONZÁLEZ-GÓMEZ, C., SERPAGLI, E., TORTELLO, M. F., VECOLI, M. and VIZCAÏNO, D. 2007. A review of the Late Cambrian (Furongian) palaeogeography in the western Mediterranean region, NW Gondwana. Earth-Science Reviews, 85, 47–81.
  • ÁLVARO, J. J., BAULUZ, B., PIERRE, C., SUBÍAS, I. and VIZCAÏNO, D. 2008. Carbon chemostratigraphy of the Cambrian-Ordovician transition in a midlatitude mixed platform, Montagne Noire, France. Geological Society of America Bulletin, 120, 962–975.
  • ARMSTRONG, H. A. 1996. Biotic recovery after mass extinction: the role of climate and ocean-state in the post-glacial (Late Ordovician–Early Silurian) recovery of the conodonts. 105–118. In HART, M. B. (ed.). Biotic recovery from mass extinction events, Vol. 102. Geological Society, Special Publication, London, 392 pp.
  • BERGERON, J. 1889. Étude géologique du massif ancien situé au Sud du Plateau Central (Thèse de la Faculté des Sciences, Paris). Annales de la Société Géologique, Paris, 22, 1–362.
  • BRETT, C. E., MOFFAT, H. E. and TAYLOR, W. L. 1997. Echinoderm taphonomy, taphofacies, and Lagersttten. Paleontological Society Papers, 3, 147–190.
  • BROCK, G. A., ENGELBRETSEN, M. J., JAGO, J. B., KRUSE, P. D., LAURIE, J. R., SHERGOLD, J. H., SHI, G. R. and SORAUF, J. E. 2000. Palaeobiogeographic affinities of Australian Cambrian faunas. Memoir of the Association of Australasian Palaeontologists, 23, 1–61.
  • BRUGUIÈRE, J. G. 1791. Tableau Encyclopédique et Méthodique des Trois Règnes de la Nature, contenant l’Helminthologie, ou les Vers Infusoires, les Vers Intestins, les Vers Mollusques, etc., Vol. 7. Panckoucke, Paris, 132 pp.
  • CABIBEL, J., TERMIER, H. and TERMIER, G. 1959. Les échinodermes mésocambriens de la Montagne Noire (Sud de la France). Annales de Paléontologie, 44, 281–294.
  • COURTESSOLE, R., PILLET, J. and VIZCAÏNO, D. 1988. Stratigraphie et paléontologie du Cambrien moyen gréseux de la Montagne Noire (versant méridional). Mémoires de la Société d’Etudes Scientifiques de l’Aude, 55 pp.
  • DESTOMBES, J. 1964. Sur l’attribution au Cambrien du Paléozoïque de Taffraout, près de Goulmina (versant méridional du Haut-Atlas central, Maroc). Comptes Rendus Sommaires de la Société Géologique de France, 5, 1–193.
  • FEIST, R. and COURTESSOLE, R. 1984. Découverte de Cambrien supérieur à trilobites de type est-asiatique dans la Montagne Noire (France méridionale). Comptes Rendus de l’Académie des Sciences, Paris (2e Série), 298, 177–182.
  • FRIEDRICH, W. P. 1993. Systematik und Funktionsmorphologie mittelkambrischer Cincta (Carpoidea, Echinodermata). Beringeria, 7, 3–190.
  • FRIEDRICH, W. P. 1995. Neue Nachweise mittelkambrischer Cincta (Carpoidea, Echinodermata) aus Marokko, Sardinien und Süd-Wales. In GEYER, G. and LANDING, E. (eds). The lower–middle Cambrian standard of western Gondwana, Beringeria, Special Issue, 2, 255–269.
  • GEYER, G. and LANDING, E. 2004. A unified Lower–Middle Cambrian chronostratigraphy for West Gondwana. Acta Geologica Polonica, 54, 179–218.
  • GEYER, G., LANDING, E. and HELDMAIER, W. 1995. Faunas and depositional environments of the Cambrian of the Moroccan Atlas regions. In GEYER, G. and LANDING, E. (eds). The lower–middle Cambrian standard of western Gondwana, Beringeria, Special Issue, 2, 47–120.
  • GIL CID, M. D. and DOMÍNGUEZ, P. 1995a. Presencia de Gyrocystis Jaekel, 1918 en el Cámbrico Medio de Zafra (Badajoz). Revista de la Sociedad Geológica de España, 8, 99–110.
  • GIL CID, M. D. and DOMÍNGUEZ, P. 1995b. Gyrocystis cruzae, una nueva especie de Cincta (Echinodermata Carpoidea) del Cambrico Medio del Ferredal de Quintana (Asturias, España). Boletín Geológico y Minero, 106, 517–531.
  • GONZÁLEZ-GÓMEZ, C. 2005. Linguliformean brachiopods of the Middle–Upper Cambrian transition from the Val d’Homs Formation, southern Montagne Noire, France. Journal of Paleontology, 79, 29–47.
  • GUTIÉRREZ-MARCO, J. C., CHAUVEL, J., MELÉNDEZ, B. and SMITH, A. B. 1984. Los equinodermos (Cystoidea, Homalozoa, Stelleroidea, Crinoidea) del Paleozoico inferior de los Montes de Toledo y Sierra Morena (España). Estudios Geológicos, 40, 421–453.
  • HARRIES, P. J., KAUFFMAN, E. G. and HANSEN, T. A. 1996. Models for biotic survival following mass extinction, 41–60. In HART, M. D. (ed.). Biotic recovery from mass extinction events, Vol. 102. Geological Society, Special Publication, London, 392 pp.
  • HERNÁNDEZ-SAMPELAYO, P. 1935. El Sistema Cambriano. Explicación del nuevo mapa geológico de España. Memorias del IGME, 1(1935), 291–518.
  • JAEKEL, O. 1918. Phylogenie und System der Pelmatozoen. Palontologische Zeitschrift, 3, 1–128.
  • JEFFERIES, R. P. S. 1990. The solute Dendrocystoides scoticus from the upper Ordovician of Scotland and the ancestry of chordates and echinoderms. Palaeontology, 33, 631–679.
  • JOSOPAIT, V. 1972. Das Kambrium und das Tremadoc von Ateca (Westliche Iberischen Ketten, NE-Spanien). Münstersche Forschungen zur Geologie und Palontologie, 23, 1–121.
  • KLEIN, J. T. 1734. Naturalis dispositio echinodermatum. Accessit lucubratiuncula de aculeis echinorum marinorum, cum spicilegio de belemnitis. Schreiber, Gedani.
  • KÜPPERS, A. N. and POHLER, S. M. L. 1992. Discovery of the first Early Ordovician conodonts from the Montagne Noire, southern France. 487–494. In WEBBY, B. D. and LAURIE, J. R. (eds). Global perspectives on Ordovician geology. A. A. Balkema, Rotterdam, 513 pp.
  • LEFEBVRE, B. 2007. Early Palaeozoic palaeobiogeography and palaeoecology of stylophoran echinoderms. Palaeogeography, Palaeoclimatology, Palaeoecology, 245, 156–199.
  • LIÑÁN, E. and GOZALO, R. 1986. Trilobites del Cámbrico Inferior y Medio de Murero (Cordillera Ibérica). Memorias del Museo Paleontológico de la Universidad de Zaragoza, 2, 1–104.
  • LIÑÁN, E., PEREJÓN, A. and SDZUY, K. 1993. The Lower–Middle Cambrian stages and stratotypes from the Iberian Peninsula: a revision. Geological Magazine, 130, 817–833.
  • LIÑÁN, E., VILLAS, E., GÁMEZ-VINTANED, J. A., ÁLVARO, J. J., GOZALO, R., PALACIOS, T. and SDZUY, K. 1996. Síntesis paleontológica del Cámbrico y Ordovícico de la Cordillera Ibérica (Cadenas Ibéricas y Cadenas Hespéricas). Revista Española de Paleontología, nº extraordinario, 21–32.
  • LIÑÁN, E., GOZALO, R., DIES, M. E., GÁMEZ VINTANED, J. A., MAYORAL, E., CHIRIVELLA, J., ESTEVE, J., ZAMORA, S., ZHURAVLEV, A. Y. and ANDRÉS, J. A. 2008. Fourth International Trilobite Conference, Trilo 08. Toledo, Spain, 2008. Post-Conference Field Trip. Lower and Middle Cambrian trilobites of selected localities in Cadenas Ibéricas (NE, Spain), Zaragoza, 52 pp.
  • MALLADA, L. and BUITRAGO, J. 1878. La fauna primordial a uno y otro lado de la Cordillera Cantábrica. Boletín de la Comisión del Mapa Geológico de España, 5, 177–194.
  • NEBELSICK, J. H. 2004. Taphonomy of Echinoderms: a review. 471–478. In HEINZELLER, T. H. and NEBELSICK, J. H. (eds). Echinoderms. Proceedings of the 11th International Echinoderm Meeting, Munich. Taylor and Francis, Rotterdam, 633 pp.
  • NORELL, M. A. 1992. Taxic origin and temporal diversity: the effect of phylogeny. 89–118. In NOVACEK, M. J. and WHEELER, Q. D. (eds). Extinction and phylogeny. Columbia University Press, New York, 252 pp.
  • PALMER, A. R. 1984. The biomere problem: evolution of an idea. Journal of Paleontology, 58, 599–611.
  • PRADO, C., DE VERNEUIL, E. and BARRANDE, J. 1860. Sur l’existence de la faune primordiale dans la Chaîne Cantabrique. Bulletin de la Société Géologique de France (2e Série), 17, 516–554.
  • RAHMAN, I. A. and ZAMORA, S. 2009. The oldest cinctan carpoid (stem-group Echinodermata), and the evolution of the water vascular system. Zoological Journal of the Linnean Society, 157, 420–432.
  • ROZHNOV, S. V. 2006. Carpozoan echinoderms from the Middle Cambrian (Mayaktakh Formation) of Siberia (lower reaches of the Lena river). Paleontological Journal, 40, 266–275.
  • SCHMITZ, U. 1971. Stratigraphie und Sedimentologie im Kambrium und Tremadoc der Westlichen Iberischen Ketten nördlich Ateca. Münster. Münstersche Forschungen zur Geologie und Palontologie, 22, 1–123.
  • SCHROEDER, R. 1973. Carpoideen aus dem Mittelkambrium Nordspaniens. Palaeontographica Abteilung A, 141, 119–142.
  • SDZUY, K. 1972. Das Kambrium der acadobaltischen Fauneprovinz. Gegenwrtiger Kenntnisstand und Probleme. Zentralblat für Geologie und Palontologie, 2, 1–91.
  • SDZUY, K. 1993. Early Cincta (Carpoidea) from the Middle Cambrian of Spain. Beringeria, 8, 189–207.
  • SHERGOLD, J. H. and SDZUY, K. 1991. Late Cambrian trilobites from the Iberian Mountains, Zaragoza Province, Spain. Beringeria, 4, 193–235.
  • SHERGOLD, J. H., FEIST, R. and VIZCAÏNO, D. 2000. Early Late Cambrian trilobites of Australo-Sinian aspect from the Montagne Noire, southern France. Palaeontology, 43, 599–632.
  • SMITH, A. B. 1988. Patterns of diversification and extinction in Early Palaeozoic echinoderms. Palaeontology, 31, 799–828.
  • SMITH, A. B. 1994. Systematics and the fossil record: documenting evolutionary patterns. Blackwell Scientific, Oxford, 223 pp.
  • SMITH, A. B. 2005. The pre-radial history of echinoderms. Geological Journal, 40, 255–280.
  • SMITH, A. B. and MCGOWAN, A. J. 2007. The shape of the Phanerozoic marine palaeodiversity curve: how much can be predicted from the sedimentary rock record of western Europe? Palaeontology, 50, 1–10.
  • SMITH, A. B. and ZAMORA, S. 2009. Rooting phylogenies of problematic fossil taxa; a case study using cinctans (stem-group echinoderms). Palaeontology, 52, 803–821.
  • TERMIER, H. and TERMIER, G. 1973. Les Echinodermes Cincta du Cambrien de la Montagne Noire. Geobios, 6, 243–266.
  • UBAGHS, G. 1967. Homostelea. S565–S581. In MOORE, R. C. (ed.). Treatise on invertebrate paleontology. Part S. Echinodermata 1 (2). Geological Society of America, Boulder, Colorado and University of Kansas Press, Lawrence, Kansas, 650 pp.
  • UBAGHS, G. 1999. Echinodermes nouveaux du Cambrien supérieur de la Montagne Noire (France méridionale). Geobios, 31, 809–829.
  • VIZCAÏNO, D. and ÁLVARO, J. J. 2003. Adequacy of the Lower Ordovician trilobite record in the southern Montagne Noire (France): biases for biodiversity documentation. Transactions of the Royal Society of Edinburgh: Earth Sciences, 93, 1–9.
  • VIZCAÏNO, D. and LEFEBVRE, B. 1999. Les echinoderms du Paléozoïque inférieur de Montagne Noire: biostratigraphie et paléodiversité. Geobios, 32, 353–364.
  • VIZCAÏNO, D., MONCERET, E., MONCERET, S. and ÁLVARO, J. J. 2008. Toward an integrated stratigraphy of the middle Cambrian–Furongian–Tremadocian from the southern Montagne Noire, France. Cuadernos del Museo Geominero, 9, 415–419.
  • WOTTE, T., ÁLVARO, J. J., SHIELDS, G. A., BRASIER, M. and VEIZER, J. 2007. C-, O- and Sr-isotope stratigraphy across the Lower-Middle Cambrian transition of the Cantabrian Mountains (Spain) and the Montagne Noire (France), West Gondwana. Palaeogeography, Palaeoclimatology, Palaeoecology, 256, 47–70.
  • ZAMORA, S. and RAHMAN, I. A. 2008. Nuevos datos sobre el género Sucocystis (Cincta, Echinodermata) en el Cámbrico Medio de España: implicaciones biostratigráficas y filogenéticas. Revista Española de Paleontología, 23, 301–313.
  • ZAMORA, S. and SMITH, A. B. 2008. A new Middle Cambrian cinctan (stem group echinoderm) from Spain convergent on stylophorans: palaeobiological implications. Acta Palaeontologica Polonica, 53, 207–220.
  • ZAMORA, S., LIÑÁN, E., GÁMEZ VINTANED, J. A., DOMÍNGUEZ ALONSO, P. and GOZALO, R. 2007. Nuevo carpoideo de la clase Cincta Jaekel, 1918 del norte de España: inferencias sobre la morfología funcional del opérculo. Ameghiniana, 44, 727–738.
  • ZAMORA, S., GOZALO, R. and LIÑÁN, E. 2009. Middle Cambrian gogiid echinoderms from Northeast Spain: taxonomy, palaeoecology and palaeogeographic implications. Acta Paleontologica Polonica, 54, 253–265.
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