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Article: New data on molluscs and their shell microstructures from the Middle Cambrian Gowers Formation, Australia

Publication: Palaeontology
Volume: 53
Part: 1
Publication Date: January 2010
Page(s): 97 – 135
Author(s): Michael J. Vendrasco, Susannah M. Porter, Artem Kouchinsky, Guoxiang Li and Christine Z. Fernandez
Addition Information

How to Cite

VENDRASCO, M. J., PORTER, S. M., KOUCHINSKY, A., LI, G., FERNANDEZ, C. Z. 2010. New data on molluscs and their shell microstructures from the Middle Cambrian Gowers Formation, Australia. Palaeontology, 53, 1, 97–135.

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References

  • ATKINS, C. J. and PEEL, J. S. 2004. New species of Yochelcionella (Mollusca; Helcionelloida) from the Lower Cambrian of North Greenland. Bulletin of the Geological Society of Denmark, 51, 1–9.
  • BENGTSON, S. 1992. Proterozoic and earliest Cambrian skeletal metazoans. 1017–1033. In SCHOPF, J. W. and KLEIN, C. (eds). The Proterozoic biosphere. Cambridge University Press, Cambridge, 1374 pp.
  • BENGTSON, S. and CONWAY MORRIS, S. 1992. Early radiation of biomineralizing taxa. 447–481. In LIPPS, J. H. and SIGNOR, P. W. (eds). Origin and evolution of the metazoa. Plenum Press, New York, 578 pp.
  • BENGTSON, S., ZHAO, Y., COOPER, B. J., JELL, P. A. and RUNNEGAR, B. N. 1990. Early Cambrian fossils from South Australia. Memoir of the Association of Australian Palaeontologists, 9, 364 pp.
  • BENGTSON, S. and ZHAO, Y. 1997. Fossilized metazoan embryos from the earliest Cambrian. Science, 277, 1645–1648.
  • BOUCHET, P., McLEAN, J. and WARÉN, A. 1983. Monoplacophorans in the North Atlantic. Oceanologica Acta, 6, 117–118.
  • BRIGGS, D. E. G. and KEAR, A. J. 1993. Fossilization of soft tissue in the laboratory. Science, 259, 1439–1442.
  • CARON, J.-B., SCHELTEMA, A., SCHANDER, C. and RUDKIN, D. 2006. A soft bodied mollusc with radula from the Middle Cambrian Burgess Shale. Nature, 442, 159–163.
  • CARTER, J. G. 1979. Comparative shell microstructure of the Mollusca, Brachiopoda and Bryozoa. In JOHARI, O. (director). Scanning electron microscopy/1979, II. Chicago Press Corporation, Chicago, 910 pp.
  • CARTER, J. G. 1980. Environmental and biological controls on bivalve shell mineralogy and microstructure. 69–113. In RHOADS, D. C. and LUTZ, R. A. (eds). Skeletal growth of aquatic organisms: biological records of environmental change. Plenum Press, New York, 750 pp.
  • CARTER, J. G. 1990. Glossary of skeletal biomineralization. 609–671. In CARTER, J. G. (ed.). Skeletal biomineralization: patterns, processes and evolutionary trends, Volume 1 . Van Nostrand Reinhold, New York, 832 pp.
  • CARTER, J. G. and CLARK, G. R. II. 1985. Classification and phylogenetic significance of molluscan shell microstructure. 50–71. In BOTTJER, D. J., HICKMAN, C. S. and WARD, P. D. (eds). Mollusks: notes for a short course. University of Tennessee Department of Geological Sciences Studies in Geology, 13 , Knoxville, Tennessee, 305 pp.
  • CARTER, J. G., BARRERA, E. and TEVESZ, M. J. S. 1998. Thermal potentiation and mineralogical evolution in the Bivalvia (Mollusca). Journal of Paleontology, 72, 991–1010.
  • CARTWRIGHT, J. H. E. and CHECA, A. G. 2007. The dynamics of nacre self assembly. Journal of the Royal Society Interface, 4, 1–14.
  • CHECA, A., ESTEBAN-DELGADO, F. J. and RODRÍGUEZ-NAVARRO, A. B. 2007. Crystallographic structure of the foliated calcite of bivalves. Journal of Structural Biology, 157, 393–402.
  • CHECA, A., RAMÍREZ, J., GONZÁLEZ-SEGURA, A. and SÁNCHEZ-NAVAS, A. 2009. Nacre and false nacre (foliated aragonite) in extant monoplacophorans (=Triblidiida: Mollusca). Naturwissenschaften, 96, 111–122.
  • COOK, P. J. 1989. Phosphate deposits of the Georgina Basin. 533–544. In NOTHOLT, A. J. G., SHELDON, R. P. and DAVIDSON, D. F. (eds). Phosphate deposits of the world, Volume 2: phosphate rock resources. Cambridge University Press, Cambridge, 566 pp.
  • CRUZ, R., WEISSMÜLLER, G. and FARINA, M. 2003. Microstructure of Monoplacophora (Mollusca) shell examined by low-voltage field emission scanning electron and atomic force microscopy. Scanning, 25, 12–18.
  • CURREY, J. D. 1977. Mechanical properties of mother of pearl in tension. Proceedings of the Royal Society of London B, 196, 443–463.
  • CURREY, J. D. 1990. Biomechanics of mineralized skeletons. 11–25. In CARTER, J. G. (ed.). Skeletal biomineralization: patterns, processes and evolutionary trends, Volume 1 . Van Nostrand Reinhold, New York, 832 pp.
  • CURREY, J. D. 1999. The design of mineralized hard tissues for their mechanical functions. The Journal of Experimental Biology, 202, 3285–3294.
  • CUVIER, G. 1797. Tableau élémentaire de l’histoire naturelle des animaux, Paris, 710 pp.
  • DONOGHUE, P. C., KOUCHINSKY, A., WALOSZEK, D., BENGTSON, S., DONG, XI-PING, VAL’KOV, A. K., CUNNINGHAM, J. A. and REPETSKI, J. E. 2006. Fossilized embryos are widespread but the record is temporally and taxonomically biased. Evolution & Development, 8, 232–238.
  • FENG, W. and SUN, W. 2003. Phosphate replicated and replaced microstructure of molluscan shells from the earliest Cambrian of China. Acta Palaentologica Polonica, 48, 21–30.
  • FENG, W., MU, X. and KOUCHINSKY, A. V. 2001. Hyolith-type microstructure in a mollusc-like fossil from the Early Cambrian of Yunnan, China. Lethaia, 34, 303–308.
  • FRAKES, L. A., FRANCIS, J. E. and SYKTUS, J. I. 1992. Climate modes of the Phanerozoic: the history of the Earth’s climate over the past 600 million years. Cambridge University Press, Cambridge, 274 pp.
  • GRÉGOIRE, C. 1961. Sur la structure submicroscopique de la conchioline associée aux prismes de coquilles de mollusques. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique: Biologie, 37, 1–34.
  • GRÉGOIRE, C. 1972. Structure of the molluscan shell. 45–102. In FLORKIN, M. and SCHEER, B. T. (eds). Chemical Zoology. Academic Press, New York, 567 pp.
  • GUBANOV, A. P. and PEEL, J. S. 1999. Oelandiella, the earliest Cambrian helcionelloid mollusc from Siberia. Palaeontology, 42, 211–222.
  • GUBANOV, A. P. and PEEL, J. S. 2000. Cambrian monoplacophoran molluscs (Class Helcionelloida). American Malacological Bulletin, 15, 139–145.
  • GUBANOV, A. P., KOUCHINSKY, A. V., PEEL, J. S. and BENGTSON, S. 2004. Middle Cambrian molluscs of ‘Australian’ aspect from northern Siberia. Alcheringa, 28, 1–20.
  • HARPER, E. M., PALMER, T. J. and ALPHEY, J. R. 1997. Evolutionary response by bivalves to changing Phanerozoic sea-water chemistry. Geological Magazine, 134, 403–407.
  • HASZPRUNAR, G. 1992. The first molluscs – small animals. Bollettino di Zoologia, 59, 1–16.
  • HAYAMI, I. 1988. Taxonomic characters of Propeamussiids from Japan. Venus, 47, 71–82.
  • HE TINGGUI and YANG XIANHE 1982. Lower Cambrian Meishucun Stage of the western Yangtze stratigraphic region and its small shelly fossils. Bulletin of the Chengdu Institute of Geology, M. R., Chinese Academy of Geological Sciences, 3, 69–95. [In Chinese, with English summary].
  • HINZ-SCHALLREUTER, I. 1997. Einsaugstutzen oder Auspuff? Das Rtsel um Yochelcionella (Mollusca, Kambrium). Geschiebekunde aktuell, 13, 105–122.
  • HORNY, R. J. 1963. On the systematic position of the cyrtonelloids (Mollusca). Časopis Národního Muzea Praze, 132, 90–94.
  • HOWARD, P. F. 1989. The D Tree phosphate deposit, Georgina Basin, Australia. 551–557. In NOTHOLT, A. J. G., SHELDON, R. P. and DAVIDSON, D. F. (eds). Phosphate deposits of the world Volume 2: phosphate rock resources. Cambridge University Press, Cambridge, 566 pp.
  • JABLONSKI, D. 1985. Molluscan development. 33–49. In BROADHEAD, T. W. (ed.). Mollusks, notes for a short course. University of Tennessee, Knoxville, Tennessee, 305 pp.
  • JACKSON, A. P., VINCENT, J. F. V. and TURNER, R. M. 1988. The mechanical design of nacre. Proceedings of the Royal Society of London B, 234, 415–440.
  • JELL, P. A. 1975. Australian Middle Cambrian eodiscoids with a review of the superfamily. Palaeontographica, Abteilung A, 150, 1–97.
  • KOBAYASHI, I. 1971. Internal shell microstructure of recent bivalvan molluscs. Science Reports of Niigata University, Series E, 2, 27–73.
  • KOUCHINSKY, A. 1999. Shell microstructures of the Early Cambrian Anabarella and Watsonella as new evidence on the origin of the Rostroconchia. Lethaia, 32, 173–180.
  • KOUCHINSKY, A. 2000. Shell microstructures in Early Cambrian molluscs. Acta Palaeontologica Polonica, 45, 119–150.
  • KOUCHINSKY, A. 2001. Mollusks, hyoliths, stenothecoids, and coeloscleritophorans. 326–349. In ZHURAVLEV, A. Y. and RIDING, R. The ecology of the Cambrian radiation. Columbia University Press, New York, 525 pp.
  • LANDING, E. and BARTOWSKI, K. E. 1996. Oldest shelly fossils from the Taconic Allochthon and Late Early Cambrian sea-levels in Eastern Laurentia. Journal of Paleontology, 70, 741–761.
  • LI, G., STEINER, M., ZHU, X., YANG, A., WANG, H. and ERDTMANN, B. D. 2007. Early Cambrian metazoan fossil record of South China: generic diversity and radiation patterns. Palaeogeography, Palaeoclimatology, Palaeoecology, 254, 229–249.
  • LOWENSTAM, H. A. and WEINER, S. 1989. On Biomineralization. Oxford University Press, Oxford, 324 pp.
  • LUCAS, J. and PRÉVÔT, L. 1985. The synthesis of apatite by bacterial activity: mechanism. Sciences Géologiques, Mémoires, 77, 83–92.
  • MacCLINTOCK, C. 1967. Shell structure of patelloid and bellerophontoid gastropods. Peabody Museum of Natural History, Yale University Bulletin, 22, 140 pp.
  • MacKINNON, D. I. 1982. Tuarangia paparua n. gen. and n. sp., a late Middle Cambrian pelecypod from New Zealand. Journal of Paleontology, 56, 589–598.
  • MacKINNON, D. I. 1985. New Zealand late Middle Cambrian molluscs and the origin of Rostroconchia and Bivalvia. Alcheringa, 9, 65–81.
  • MARIN, F., LUQUET, G., MARIE, B. and MEDAKOVIC, D. 2008. Molluscan shell proteins: primary structure, origin, and evolution. Current Topics in Developmental Biology, 80, 209–276.
  • MARTÍ MUS, M., PALACIOS, T. and JENSEN, S. 2008. Size of the earliest mollusks: did small helcionellids grow to become large adults? Geology, 36, 175–178.
  • McLEAN, J. 1979. A new monoplacophoran limpet from continental shelf off southern California. Contributions in Science, Natural History Museum of Los Angeles County, 307, 1–19.
  • MISSARZHEVSKY, V. V. and MAMBETOV, A. M. 1981. Stratigraphy and fauna of the Precambrian-Cambrian boundary beds of Malyj Karatau. Trudy Geologiske Institut Leningrad, 326, 1–96.
  • MÜLLER, K. J. and HINZ-SCHALLREUTER, I. 1993. Palaeoscolecid worms from the Middle Cambrian of Australia. Palaeontology, 36, 549–592.
  • MUTVEI, H. 1980. The nacreous layer in molluscan shells. 49–56. In OMORI, M. and WATABE, N. (eds). The mechanisms of biomineralization in animals and plants. Tokai University Press, Tokyo, 310 pp.
  • MUTVEI, H. 1984. Ultrastructural research on molluscan nacre. Proceedings of the 27th International Geological Congress, 2, 111–124.
  • ÖPIK, A. A. 1979. Middle Cambrian agnostids: systematics and biostratigraphy. Bureau of Mineral Resources Bulletin (Geology and Geophysics, Australia), 172, 188 pp.
  • PARKHAEV, P. Yu. 2001a. Trenella bifrons gen. et sp. nov.: a new helcionelloid mollusk from the Lower Cambrian of South Australia. Paleontological Journal, 35, 22–24.
  • PARKHAEV, P. YU. 2001b. Molluscs and siphonoconchs. 133–210. In ALEXANDER, E. M., JAGO, J. B., ROZANOV, A. Yu. and ZHURAVLEV, A. Yu. The Cambrian biostratigraphy of the Stansbury Basin, South Australia. Transactions of the Palaeontological Institute, 282 . Interperiodica, Moscow, 215 pp.
  • PEEL, J. S. 1980. A new Silurian retractile monoplacophoran and the origin of the gastropods. Proceedings of the Geological Association. Interperiodica, Moscow, 91, 91–97.
  • PEEL, J. S. 1991a. Functional morphology of the Class Helcionelloida nov., and the early evolution of the Mollusca. 157–177. In SIMONETTA, A. M. and CONWAY MORRIS, S. (eds). The early evolution of Metazoa and the significance of problematic taxa. Cambridge University Press, Cambridge, 308 pp.
  • PEEL, J. S. 1991b. The Classes Tergomya and Helcionelloida and early molluscan evolution. Grønlands Geologiske Undersøgelse Bulletin, 161, 11–65.
  • PEEL, J. S. 2004. Pinnocaris and the origin of scaphopods. Acta Palaeontologica Polonica, 49, 543–550.
  • PEEL, J. S. 2006. Scaphopodization in Palaeozoic molluscs. Palaeontology, 49, 1357–1364.
  • PEI, F. 1985. First discovery of Yochelcionella from the Lower Cambrian of China and its significance. Acta Micropalaeontologica Sinica, 2, 395–400.
  • PENG, S. and ROBISON, R. A. 2000. Agnostid biostratigraphy across the Middle Upper Cambrian boundary in Hunan, China. Journal of Paleontology, 74 (supplement to no. 4), 1–104.
  • PÉREZ-HUERTA, A. and CUSACK, M. 2008. Common crystal nucleation mechanism in shell formation of two morphologically distinct calcite brachiopods. Zoology, 111, 9–15.
  • POJETA, J. Jr and RUNNEGAR, B. 1976. The paleontology of rostroconch mollusks and the early history of the Phylum Mollusca. Geological Survey Professional Paper, 968, 1–88.
  • POJETA, J. JR, RUNNEGAR, B., MORRIS, N. J. and NEWELL, N. D. 1972. Rostroconchia: a new class of bivalved mollusks. Science, 177, 264–267.
  • POJETA, J. JR, GILBERT-TOMLINSON, J. and SHERGOLD, J. H. 1977. Cambrian and Ordovician rostroconch molluscs from northern Australia. Bureau of Mineral Resources, Geology and Geophysics, Australia, 171, 54 pp.
  • PORTER, S. M. 2004. Halkieriids in Middle Cambrian phosphatic limestones from Australia. Journal of Paleontology, 78, 574–590.
  • PORTER, S. M. 2007. Seawater chemistry and early carbonate biomineralization. Science, 316, 1302.
  • PRÉVÔT, L. and LUCAS, J. 1990. Phosphate. 256–257. In BRIGGS, D. E. G. and CROWTHER, P. R. (eds). Palaeobiology: a synthesis. Blackwell Scientific Publications, Oxford, 583 pp.
  • RASBAND, W. S. 19972007. ImageJ. U. S. National Institutes of Health, Bethesda, Maryland, USA, http://rsb.info.nih.gov/ij/ .
  • ROGALLA, N. S., CARTER, J. G. and POJETA, J. Jr 2003. Shell microstructure of the Late Carboniferous rostroconch mollusc Apotocardium lanterna (Branson, 1965). Journal of Paleontology, 77, 655–673.
  • ROZANOV, A. Yu., MISSARZHEVSKY, V. V., VOLKOVA, N. A., VORONOVA, L. G., KRYLOV, I. N., KELLER, B. M., KOROLYUK, I. K., LENDZION, K., MICHNIAK, R., PYKHOVA, N. G. and SIDOROV, A. D. 1969. The Tommotian Stage and the problem of the lower boundary of the Cambrian. Trudy Geologicheskogo Instituta AN SSSR, 296, 380 pp.
  • RUNNEGAR, B. 1978. Origin and evolution of the Class Rostroconchia. Philosophical Transactions of the Royal Society of London B, 284, 319–333.
  • RUNNEGAR, B. 1981. Muscle scars, shell form and torsion in Cambrian and Ordovician univalved molluscs. Lethaia, 14, 311–322.
  • RUNNEGAR, B. 1983. Molluscan phylogeny revisited. Memoirs of the Association of Australian Palaeontologists, 1, 121–144.
  • RUNNEGAR, B. 1984. Crystallography of the foliated calcite shell layers of bivalve molluscs. Alcheringa, 8, 273–290.
  • RUNNEGAR, B. 1985. Shell microstructures of Cambrian molluscs replicated by phosphate. Alcheringa, 9, 245–257.
  • RUNNEGAR, B. 1987. Class Monoplacophora. 297–304. In BOARDMAN, R. S., CHEETHAM, A. H. and ROWELL, A. J. (eds). Fossil invertebrates. Blackwell Scientific Publications, Palo Alto, California, 713 pp.
  • RUNNEGAR, B. 1989. The evolution of mineral skeletons. 75–94. In CRICK, R. E. (ed.). Origin, evolution, and modern aspects of biomineralization in plants and animals. Plenum Press, New York, 536 pp.
  • RUNNEGAR, B. 1996. Early evolution of the Mollusca: the fossil record. 77–87. In TAYLOR, J. (ed.). Origin and evolutionary radiation of the Mollusca. Oxford University Press, 408 pp.
  • RUNNEGAR, B. and BENTLEY, C. 1983. Anatomy, ecology and affinities of the Australian Early Cambrian bivalve Pojetaia runnegari Jell. Journal of Paleontology, 57, 73–92.
  • RUNNEGAR, B. and JELL, P. A. 1976. Australian Middle Cambrian molluscs and their bearing on early molluscan evolution. Alcheringa, 1, 109–138.
  • RUNNEGAR, B. and JELL, P. A. 1980. Australian Middle Cambrian molluscs: corrections and additions. Alcheringa, 4, 111–113.
  • RUNNEGAR, B. and POJETA, J. Jr 1974. Molluscan phylogeny: the paleontological viewpoint. Science, 186, 311–317.
  • RUNNEGAR, B. and POJETA, J. JR 1980. The monoplacophoran mollusc Yochelcionella identified from the Lower Cambrian of Pennsylvania. Journal of Paleontology, 54, 635–636.
  • RUNNEGAR, B. and POJETA, J. JR 1985. Origin and diversification of the Mollusca. 1–57. In TRUEMAN, E. R. and CLARKE, M. R. (eds). The Mollusca, Volume 10: Evolution. Academic Press, Orlando, Florida, 495 pp.
  • RUNNEGAR, B. and POJETA, J. JR 1992. The earliest bivalves and their Ordovician descendents. American Malacological Bulletin, 9, 117–122.
  • SAGEMANN, J., BALE, S. J., BRIGGS, D. E. G. and PARKES, R. J. 1999. Controls on the formation of authigenic minerals in association with decaying organic matter: an experimental approach. Geochimica et Cosmochimica Acta, 63, 1083–1095.
  • SHARPE, D. 1853. Description of the new species of Zoophyta and Mollusca (appendix B to Ribeiro, Carlos, Carboniferous and Silurian formations of the neighborhood of Bussaco in Portugal. Geological Society of London Quarterly Journal, 9, 146–158.
  • SHERGOLD, J. H. and LAURIE, J. R. 1986. Appendix 5: palaeontological determinations from selected formations and field localities. 80–89. In SHERGOLD, J. H. and SOUTHGATE, P. N. Middle Cambrian phosphatic and calcareous lithofacies along the eastern margin of the Georgina Basin, Western Queensland. Australasian sedimentologists group field guide series, 2, 89 pp.
  • SHERGOLD, J. H. and SOUTHGATE, P. N. 1986. Middle Cambrian phosphatic and calcareous lithofacies along the eastern margin of the Georgina Basin, Western Queensland. Australasian sedimentologists group field guide series, 2, 89 pp.
  • SKOVSTED, C. B. 2004. Mollusc fauna of the Early Cambrian Bastion Formation of North-East Greenland. Bulletin of the Geological Society of Denmark, 51, 11–37.
  • SMITH, A. G. 2001. Paleomagnetically and tectonically based global maps for Vendian to Mid-Ordovician time. 11–46. In ZHURAVLEV, A. Y. and RIDING, R. The ecology of the Cambrian radiation. Columbia University Press, New York, 525 pp.
  • SMITH, B. L., SCHÄFFER, T. E., VIANI, M., THOMPSON, J. B., FREDERICK, N. A., KINDT, J., BELCHER, A., STUCKY, G. D., MORSE, D. E. and HANSMA, P. K. 1999. Molecular mechanistic origin of the toughness of natural adhesives, fibres, and composites. Nature, 399, 761–763.
  • SOUTHGATE, P. N. 1986a. The Gowers Formation and Bronco Stromatolith Bed, two new stratigraphic units in the Undilla portion of the Georgina Basin. Queensland Government Mining Journal, October, 407–411.
  • SOUTHGATE, P. N. 1986b. Proterozoic and Cambrian phosphorites – specialist studies: Middle Cambrian phosphatic hardgrounds, phoscrete profiles and stromatolites and their implications for phosphogenesis. 327–351. In COOK, P. J. and SHERGOLD, J. H. (eds). Phosphate deposits of the world Volume 1: Proterozoic and Cambrian phosphorites. Cambridge University Press, 386 pp.
  • SOUTHGATE, P. N. and SHERGOLD, J. H. 1991. Application of sequence stratigraphic concepts to Middle Cambrian phosphogenesis, Georgina Basin, Australia. Journal of Australian Geology and Geophysics, 12, 119–144.
  • TAYLOR, J. D. and LAYMAN, M. 1972. The mechanical properties of bivalve (Mollusca) shell structures. Palaeontology, 15, 73–87.
  • TAYLOR, J. D., KENNEDY, W. J. and HALL, A. 1969. The shell structure and mineralogy of the Bivalvia: introduction, Nuculacea – Trigonacea. Bulletin of the British Museum of Natural History, Zoology, Supplement, 3, 125 pp.
  • TAYLOR, P. D. and WEEDON, M. J. 2000. Skeletal ultrastructure and phylogeny of cyclostome bryozoans. Zoological Journal of the Linnean Society, 128, 337–399.
  • USHATINSKAYA, G. T. and PARKHAEV, P. YU. 2005. Preservation of imprints and casts of cells of the outer mantle epithelium in the shells of Cambrian brachiopods, mollusks, and problematics. Palaeontological Journal, 39, 251–263.
  • WADA, K. 1961. Crystal growth of molluscan shells. Bulletin of the National Pearl Research Laboratory, Japan, 7, 703–828.
  • WADA, K. 1966. Spiral growth of nacre. Nature, 211, 1427.
  • WADA, K. 1972. Nucleation and growth of aragonite crystals in the nacre of some bivalve molluscs. Biomineralisation, 6, 141–159.
  • WAGNER, P. J. 1997. Patterns of morphologic diversification among the Rostroconchia. Paleobiology, 23, 115–150.
  • WAGNER, P. J. 2002. Phylogenetic relationships of the earliest anisotrophically coiled gastropods. Smithsonian Contributions to Paleobiology, 88, 152 pp.
  • WAHLMAN, G. P. 1992. Middle and Upper Ordovician symmetrical univalved mollusks (Monoplacophora and Bellerophontina) of the Cincinnati Arch region. U. S. Geological Survey Professional Paper, 1066–O, 213 pp.
  • WALOSSEK, D., HINZ-SCHALLREUTER, I., SHERGOLD, J. H. and MÜLLER, K. J. 1993. Three-dimensional preservation of arthropod integument from the Middle Cambrian of Australia. Lethaia, 26, 7–15.
  • WARÉN, A. and BOUCHET, P. 1990. Laevipilina rolani, a new monoplacophoran from off southwestern Europe. Journal of Molluscan Studies, 56, 449–453.
  • WARÉN, A. and GOFAS, S. 1996. A new species of monoplacophora, redescription of the genera Veleropilina and Rokopella, and new information on three species of the class. Zoologica Scripta, 25, 215–232.
  • WARÉN, A. and HAIN, S. 1992. Laevipilina antarctica and Micropilina arntzi, two new monoplacophorans from the Antarctic. The Veliger, 35, 165–176.
  • WATABE, N. 1988. Shell structure. 69–104. In TRUEMAN, E. R. and CLARKE, M. R. (eds). The Mollusca, Volume 11: form and function. Academic Press, San Diego, California, 504 pp.
  • WEBERS, G. F., POJETA, J. Jr and YOCHELSON, E. L. 1992. Cambrian molluscs from the Minaret Formation, Ellsworth Mountains, West Antarctica. Memoirs of the Geological Society of America, 170, 181–248.
  • WEEDON, M. J. and TAYLOR, P. D. 1995. Calcitic nacreous ultrastructures in bryozoans: implications for comparative biomineralization of lophophorates and molluscs. Biological Bulletin, 188, 281–292.
  • WENZ, W. 1938. Gastropoda. 1–240. In SCHINDEWOLF, O. H. (ed). Handbuch der Palozoologie, Band 6, Lief. 1. Gebrüder Borntraeger, Berlin, 480 pp.
  • WENZ, W. 1940. Ursprung und fruhe Stammesgeschichte der Gastropoden. Archiv fur Molluskenkunde, 72, 1–10.
  • WEST, K. and COHEN, A. 1996. Shell microstructure of gastropods from Lake Tanganyika, Africa: adaptation, convergent evolution, and escalation. Evolution, 50, 672–681.
  • WILLIAMS, A. 1997. Shell structure. 267–320. In KAISER, R. L. (ed.). Treatise on Invertebrate Paleontology, Part H: Brachiopoda, revised, Volume 1: introduction. Geological Society of America, Boulder, Colorado and University of Kansas Press, Kansas, 560 pp.
  • WILLIAMS, A. and WRIGHT, A. D. 1970. Shell structure of the Craniacea and other calcareous inarticulate brachiopods. Special Papers in Palaeontology, 7, 1–51.
  • XIAO, S. and KNOLL, A. H. 2000. Phosphatized animal embryos from the Neoproterozoic Doushantuo Formation at Weng’an, Guizhou, South China. Journal of Paleontology, 74, 767–788.
  • ZHOU, B. and XIAO, L. 1984. Early Cambrian monoplacophorans and gastropods from Huainan and Huoqiu counties, Anhui Province. Professional Papers of Stratigraphy and Palaeontology, Chinese Academy of Geological Sciences, 13, 125–140.
  • ZHURAVLEV, A. YU. 1993. Early Cambrian steps of biomineralization: mineralogy. Biomineralization, 93, 102 [Seventh International Symposium on Biomineralization, Monaco, Program and Abstracts].
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