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Article: Extending the footprint record of Pareiasauromorpha to the Cisuralian: earlier appearance and wider palaeobiogeography of the group

Papers in Palaeontology - Volume 7 Issue 1 - Cover
Publication: Papers in Palaeontology
Volume: 7
Part: 3
Publication Date: August 2021
Page(s): 1297 1319
Author(s): Lorenzo Marchetti, Sebastian Voigt, Eudald Mujal, Spencer G. Lucas, Heitor Francischini, Josep Fortuny, and Vincent L. Santucci
DOI: 10.1002/spp2.1342
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How to Cite

MARCHETTI, L., VOIGT, S., MUJAL, E., LUCAS, S.G., FRANCISCHINI, H., FORTUNY, J., SANTUCCI, V.L. 2021. . Papers in Palaeontology, 7, 3, 1297-1319. DOI: /doi/10.1002/spp2.1342

Author Information

  • Lorenzo Marchetti - Museum für Naturkunde, Leibniz-Institut für Evolutions-und Biodiversitätsforschung Invalidenstrasse 43 10115 Berlin Germany
  • Sebastian Voigt - Urweltmuseum Geoskop Burgstraße 19 66871 Thallichtenberg Germany
  • Eudald Mujal - Staatliches Museum für Naturkunde Stuttgart Rosenstein 1 70191 Stuttgart Germany
  • Eudald Mujal - Institut Català de Paleontologia Miquel Crusafont, Universitat Autònoma de Barcelona Universitat Autònoma de Barcelona Edifici ICTA-ICP, c/Columnes s/n, Campus de la UAB 08193 Cerdanyola del Vallès Barcelona, Spain
  • Spencer G. Lucas - New Mexico Museum of Natural History & Science 1801 Mountain Road NW Albuquerque NM 87104 USA
  • Heitor Francischini - Laboratório de Paleontologia de Vertebrados Instituto de Geociências Universidade Federal do Rio Grande do Sul Porto Alegre RS Brazil
  • Josep Fortuny - Institut Català de Paleontologia Miquel Crusafont, Universitat Autònoma de Barcelona Universitat Autònoma de Barcelona Edifici ICTA-ICP, c/Columnes s/n, Campus de la UAB 08193 Cerdanyola del Vallès Barcelona, Spain
  • Vincent L. Santucci - Geologic Resources Division National Park Service 1849 C Street, NW, Room 2644 Washington DC 20240 USA

Publication History

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    Pareiasauromorpha is one of the most important tetrapod groups of the Permian. Skeletal evidence suggests a late Kungurian origin in North America, whereas the majority of occurrences come from the Guadalupian and Lopingian of South Africa and Russia. However, Pareiasauromorpha footprints include the ichnogenus Pachypes, which is unknown from strata older than late Guadalupian. A revision of several Pachypes-like footprints from the Cisuralian–Guadalupian of Europe and North America confirm the occurrence of this ichnogenus and of the ichnospecies Pachypes ollieri comb. nov. beginning in the Artinskian. This is the earliest known occurrence of Pachypes and it coincides with the Artinskian reptile radiation. Based on a synapomorphy-based track–trackmaker correlation, P. ollieri can be attributed to nycteroleter pareiasauromorphs such as Macroleter. Therefore, the earliest occurrences of pareiasauromorph footprints precede by at least 10 myr the earliest occurrence of this group in the skeletal record. Moreover, the palaeobiogeography of the group is extended to the Cisuralian and Guadalupian of western Europe.


    For access to collections/sites and other assistance during the study we warmly thank: Bernard Mourgues of the Société de Protection de la Nature Languedoc Roussillon, Stéphane Fouché of the Musée Fleury, Laurent Marivaux, Pierre-Olivier Antoine, Maeva Orliac, Monique Vianey-Liaud and Suzanne Jiquel of the Institut des Sciences de l’Évolution de Montpellier, University of Montpellier, Nour-Eddine Jalil of the Muséum National d'Histoire Naturelle, Anne Joncheray of the Musée Archéologique de Saint-Raphaël, Emmanuel Fara and Jerome Thomas of the University of Dijon, Joanne Turner and Fiona Wilson from the Dumfries Museum and Camera Obscura, Maria Rita Palombo and Linda Riti of the Palaeontological Museum of the ‘La Sapienza’ University of Rome, Robert Reisz of the University of Toronto at Mississauga and Valeriy V. Bulanov, Valeriy K. Golubev, and Yuri M. Gubin of the Paleontological Institute of Moscow. We extend our thanks to Colleen Hyde, Kim Besom and Klara Widrig, Grand Canyon National Park, for their assistance obtaining photographs of the track specimen from the Hermit Shale in the park collection. We acknowledge Sergio Llàcer, Institut Català de Paleontologia (ICP), for his assistance in the creation of several 3D photogrammetric models. We would like to thank Lara Sciscio, Massimo Bernardi, Marc J. Van den Brandt and an anonymous reviewer for useful suggestions, as well as the editors of the journal. LM was supported by the Alexander von Humboldt Foundation for his postdoc at the Urweltmuseum Geoskop of Thallichtenberg. LM is funded by the Bundesministerium für Bildung und Forschung (BMBF) for the BROMACKER project 2020. EM received funding for collection visits from: the SYNTHESYS Project (FR-TAF-3621, FR-TAF-4808; which is financed by European Community Research Infrastructure Action under the FP7 ‘Capacities’ Program, Secretaria d'Universitats i de Recerca (Departament d'Economia i Coneixement, Generalitat de Catalunya; exp. num. 2013 CTP 00013) and the Erasmus+ program (2016) of the Universitat Autònoma de Barcelona. JF is supported by the Spanish Agencia Estatal de Investigación and the European Regional Development Fund of the European Union (AEI/FEDER EU, project CGL2017-82654-P). JF is member of the consolidated research group 2017 SGR 86 GRC of the Generalitat de Catalunya. We acknowledge support from the CERCA programme (ICP) from the Generalitat de Catalunya, and the project ‘Evolució dels ecosistemes durant la transició Paleozoic–Mesozoic a Catalunya’ (ref. CLT009/18/00066), financially supported by the Departament de Cultura (Generalitat de Catalunya).

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