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Article: New species of Franchia and Protozigzagiceras (Ammonoidea, Middle Jurassic): the phyletic origin of Zigzagiceratinae

Publication: Palaeontology
Volume: 57
Part: 4
Publication Date: July 2014
Page(s): 713 741
Author(s): <p>Sixto R. Fernandez-Lopez and Giulio Pavia</p>
Addition Information

How to Cite

FERNANDEZ-LOPEZ, S.R., PAVIA, G. 2014, New species of Franchia and Protozigzagiceras (Ammonoidea, Middle Jurassic): the phyletic origin of Zigzagiceratinae. Palaeontology, 57, 4, 713–741. doi: 10.1111/pala.12084

Author Information

  • Sixto R. Fernandez-Lopez - Departamento de Paleontología, Facultad de Ciencias Geológicas, Universidad Complutense de Madrid, Madrid, Spain (email:
  • Giulio Pavia - Dipartimento di Scienze della Terra, Torino, Italy (email:

Publication History

  • Issue published online: 14 JUL 2014
  • Article first published online: 18 NOV 2013
  • Manuscript Accepted: 28 SEP 2013
  • Manuscript Received: 10 MAY 2012

Funded By

MICINN. Grant Numbers: CGL2008-01273, CGL2011-23947
Torino University Grants 2008

Online Version Hosted By

Wiley Online Library
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Three genera and seven species belonging to the subfamily Zigzagiceratinae (family Perisphinctidae) are described from the Lower Bathonian of France and Saudi Arabia. Intraspecific dimorphism is recognized. A revision of the genus Franchia proposed by Sturani (1967), based on the syntypes and new specimens from south-east France, is presented. Franchia arkelli Sturani, Franchia subalpina sp. nov., Protozigzagiceras torrensi (Sturani), Protozigzagiceras tethycum sp. nov., Protozigzagiceras flexum sp. nov. and Protozigzagiceras densum sp. nov. are described from the Digne–Castellane region of south-east France. Megazigzagiceras subarabicum, gen. et sp. nov. is described from the Dharma region of Saudi Arabia. The successive Early Bathonian species of Franchia and Protozigzagiceras herein identified in West Tethyan areas, as members of the Mediterranean–Caucasian Subrealm, formed lasting separate peramorphoclines characterized by increasing hydrodynamic coiling of the shell. In contrast, rapid proterogenesis originated and diversified the earliest Bathonian zigzagiceratin lineages, giving paedomorphic members, commonly neotenic and more scarcely progenetic. ProceritesSiemiradzkia seems to be the oldest zigzagiceratin member in the French Subalpine, Iberian and Lusitanian basins, branched off by paedomorphosis from leptosphinctins at the Bajocian–Bathonian transition. The Mediterranean–Caucasian genera Franchia, Zigzagiceras, Zigzagites and Wagnericeras branched from successive species of Protozigzagiceras, in turn, a direct derivative of Procerites. The oldest lineages of the clade Zigzagiceratinae evolved by iterative, rapid, paedomorphic changes and additional, lasting, peramorphic modifications during the Early Bathonian.

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