Article: A new genus of calceocrinid from Spain with comments on mosaic evolution
Espanocrinus lemonei n. gen., n. sp. is the first calceocrinid crinoid reported from the Early Devonian of Spain. This genus has primitive main axil and arm structure and advanced dorsal cup features. This suggests that mosaic evolution occurred in the calceocrinids and that cup changes were most significant in Ordovician to Devonian time. Modifications of the main axil and arm were thereafter most important in some lineages.