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Article: Taxonomy of the proterosuchid archosauriforms (Diapsida: Archosauromorpha) from the earliest Triassic of South Africa, and implications for the early archosauriform radiation

Publication: Palaeontology
Volume: 58
Part: 1
Publication Date: January 2015
Page(s): 141 170
Author(s): Martín D. Ezcurra and Richard J. Butler
Addition Information

How to Cite

EZCURRA, M.D., BUTLER, R.J. 2015. Taxonomy of the proterosuchid archosauriforms (Diapsida: Archosauromorpha) from the earliest Triassic of South Africa, and implications for the early archosauriform radiation. Palaeontology, 58, 1, 141-170.

Author Information

  • Martín D. Ezcurr - School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, UK (email: martindezcurra@yahoo.com.ar)
  • Martín D. Ezcurr - GeoBio-Center, Ludwig-Maximilians-Universität München, Munich, Germany
  • Richard J. Butler - School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, UK (email: r.butler.1@bham.ac.uk)
  • Richard J. Butler - GeoBio-Center, Ludwig-Maximilians-Universität München, Munich, Germany

Publication History

  • Issue published online: 7 JAN 2015
  • Article first published online: 20 OCT 2014
  • Manuscript Accepted: 29 AUG 2014
  • Manuscript Received: 30 JUN 2014

Funded By

DFG Emmy Noether Programme. Grant Number: BU 2587/3-1
Marie Curie Career Integration Grant. Grant Number: PCIG14-GA-2013-630123

Online Version Hosted By

Wiley Online Library (Free Access)
Get Article: Wiley Online Library [Free Access]

Abstract

Proterosuchidae is one of the first clades of Archosauriformes (archosaurs and closely related species) to appear in the fossil record, with the richest sample of the group coming from the Lystrosaurus Assemblage Zone (earliest Triassic) of South Africa. Four nominal proterosuchid species were described from South Africa during the twentieth century (Proterosuchus fergusi, Chasmatosaurus vanhoepeni, Chasmatosaurus alexanderi and Elaphrosuchus rubidgei), but interpretations of their taxonomy have been widely disparate. The most recent taxonomic revision concluded that P. fergusi is the only valid species and that the other nominal species are junior subjective synonyms of this taxon. This proposal was based on the interpretation that anatomical differences between the nominal species could be explained as a result of ontogenetic changes and/or post-mortem deformation. The recent discoveries of multiple new South African proterosuchid specimens provide an impetus to revisit their taxonomy. Based upon a comprehensive re-examination of all known specimens, as well as examination of other proterosuchid taxa in collections worldwide, we conclude that the holotype of Proterosuchus fergusi is undiagnostic. As a result, we propose a neotype (RC 846) for the species. ‘Chasmatosaurus vanhoepeni’ and ‘Elaphrosuchus rubidgei’ are considered subjective junior synonyms of P. fergusi. ‘Chasmatosaurusalexanderi is considered a valid species, for which we propose the new combination P. alexanderi comb. nov. A third species, P. goweri sp. nov., is erected on the basis of a single specimen (NMQR 880). All three species recognized here are taxonomically distinct from a previously described archosauriform maxilla from the lower Lystrosaurus AZ. As a result, we recognize a minimum of four archosauriform species following the Permo-Triassic mass extinction in South Africa. Our results suggest a greater species richness of earliest Triassic archosauriforms than previously appreciated, but that archosauriform morphological disparity remained low and did not expand until the late Early Triassic – early Mid-Triassic.

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